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More recently, increased understanding of nuclease function has led to more direct DNA editing, using techniques such as zinc finger nucleases and CRISPR. The expressed nucleases then knock out and replace genes in the chromosome.As of 2014 these approaches involve removing cells from patients, editing a chromosome and returning the transformed cells to patients.The first therapeutic use of gene transfer as well as the first direct insertion of human DNA into the nuclear genome was performed by French Anderson in a trial starting in September 1990.Between 1989 and February 2016, over 2,300 clinical trials had been conducted, more than half of them in phase I.

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The target DNA is then analyzed for the presence of the probe via radioactivity or fluorescence.

In this experiment, as in most molecular biology techniques, a control must be used to ensure successful experimentation.

Short (20-25 nucleotides in length), labeled probes are exposed to the non-fragmented target DNA, hybridization occurs with high specificity due to the short length of the probes and even a single base change will hinder hybridization.

The target DNA is then washed and the labeled probes that didn't hybridize are removed.

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